A direct continuation of the former season, this series again follows the story of Nozomi Yumehara and her friends from Yes! Precure 5. The girls who had formerly lost their powers and bade farewell to their friends Coco and Nuts from Palmier Kingdom are resurrected as Pretty Cure by the mysterious woman Flora who also wants them to find her in a place called Cure Rosegarden. To go there, they need the magic Rose Pact and the powers of the four kings of countries surrounding Palmier. But a dark organization called Eternal is also striving to get to Cure Rosegarden by stealing the Rose Pact. Luckily, Precure get helped by new allies: the delivery-boy Syrup who can change into a giant bird and fly them anywhere they want and the mysterious and beautiful fighter Milky Rose stand alongside Pretty Cure to protect what is really valuable.

Extra anime shorts included on DVD. Episode 1: Summer Memories (Vol. 1) Episode 2: The Meteor Distance (Vol. 3) Episode 3: Twinkle Star (Vol. 5) Note: Music video specials are not included in the entry as they don't contain any new scenes.

Meg, faithful and gentle, Jo, independent and energetic, Beth, quiet and shy, Amy, amiable and precocious–the four sisters live vigorously and happily at home with their gentle mother while waiting for their father to return from the Civil War. Helping each other, they challenge their fates and grow up to become wonderful women.

Happy-go-lucky short stories of daily lives of the band members of Plasmagica and ShinganCrimsonZ.